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Each PhD Program offers its PhD students a specialized set of introductory and advanced courses as well as symposia and retreats.
At the institutionary level, there are a number of specialized courses e.g. on microscopy, laboratory animal science, etc. on offer as well.
For details, please check the websites of the individual programs and institutions.
PhD Programs
Biomedical Ethics and Law - Medical Track
Biomolecular Structure and Mechanism
Epidemiology and Biostatistics
Integrative Molecular Medicine